Generically, it is the coefficient of the term linear in the gauge field in a gauge-invariant action, so it is the "matter stuff" a gauge field propagator abuts to in a Feynman diagram. Classically, it corresponds to a conserved global-invariance Noether current associated to a conserved charge.
In the gauge theory, it is the linchpin of gauge-invariance and covariant derivative completion. QFT texts focus on describing the global-to-local symmetry conversion elegantly, but leave the obvious stuff implied. I will illustrate the obvious stuff with U(1), QED, and a complex scalar field (since it contains an extra ingredient (seagulls) lacking in the spinor case), and then only cursorily extend to the non-abelian case, differing only in group theoretical aspects, covered amply in the texts. I may be cavalier with some coefficients.
$$
{\cal L}_{matter}= \tfrac{1}{2} \partial_\mu \phi^* \partial^\mu \phi + ... ,
$$
where the ellipsis ... indicates mass and interaction terms without derivatives, important in the E-L eqns of motion, but irrelevant to the gauge invariance we'll be discussing. This action is invariant under $\phi \mapsto e^{i\theta} \phi$, for constant angle θ, but not for spacetime dependent θ(x). The Noether current of the global symmetry is
$$
J^\mu_N= \tfrac{i}{2} (\phi \partial^\mu \phi^* -\phi^* \partial^\mu \phi ), \tag{1}
$$
conserved on-shell, i.e., by use of the E-L eqns of motion (which also involve the omitted non-derivative terms).
However, for local θ(x), there are extra uncancelled pieces in the transformation,
$$
\delta {\cal L}_{matter}=\partial^\mu \theta ~ J^N_\mu + \partial_\mu \theta \partial^\mu\theta ~ \phi^* \phi. \tag{2}
$$
Still, these can be cancelled by first adding a gauge action
$$
{\cal L}_{gauge}= -\tfrac{1}{4} F_{\mu\nu} F^{\mu\nu},
$$
where $A_\mu\mapsto A_\mu + \tfrac{1}{e} \partial_\mu \theta$, rendering it locally invariant, and also coupling terms cancelling (2),
$$
{\cal L}_{int}= -e A^\mu j^N_\mu + \frac{e^2}{2} A_\mu A^\mu \phi^* \phi\\
=-\frac {i e}{2} A^\mu \left ( \phi (\partial_\mu +ieA_\mu )\phi^* -\phi^* (\partial_\mu -ieA_\mu )\phi \right ) , \tag{3}
$$
rendering
$$
{\cal L}_{matter} + {\cal L}_{int} = \tfrac{1}{2} \left((\partial_\mu -ieA_\mu) \phi\right )^* (\partial^\mu -ieA^\mu) \phi + ... ,
$$
separately gauge invariant. (Off-shell: one never needs to know about the E-L equations.) A fortiori, it is globally U(1) invariant, with the gauge field not transforming here. (In the non-abelian case, it only needs transform like a matter field.)
Now, note $J^N_\mu$ is not gauge invariant—it needs the bilinear ("seagull") term in (3) to cancel its gauge variation. People usually call it the symmetry current, and, as you saw, the easiest way to isolate it is to look for the coefficient of
$\partial_\mu \theta$ in a gauge variation of the action.
Diagrammatically, it is what the gauge boson propagator ends at, coupling to φ and φ*. The seagull term has two photons emanating from it, like the wings of a seagull.
As you see from the second line of (3), you may combine everything into a gauge-invariant current involving the gauge field as well ($J_\mu^N-eA_\mu \phi\phi^*$, the Noether current of the full gauge action). You may check that, now, with the full gauge action, it is this gauge-invariant current that is conserved on-shell, while the original $J^N_\mu$ isn't, anymore, by itself. The lagrangian has changed.
Further, note the current need not be bilinear in the matter fields; for fancy interactions like
$${\cal L}_{matter}= \tfrac{f(\phi^* \phi)}{2} \partial_\mu \phi^* \partial^\mu \phi $$
you would have, instead
$$
J^\mu_N= \tfrac{if(\phi^* \phi)}{2} (\phi \partial^\mu \phi^* -\phi^* \partial^\mu \phi ), $$
for all kinds of baroque functions f(|φ|2). For SSB systems, you'd have the current be proportional to the gradient of the Goldstone boson and less significant higher-order terms in the fields.
- The extension to the non-Abelian case is straightforward, and basically a test of your appreciation of the group theoretical notation, involving group generator matrices $T_a$, group transformations such as $U=e^{i\theta ^aT_a}$, traces in their spaces, and group covariant derivatives,
$$
J^a_\mu = i (\partial_\mu \Phi^\dagger T^a \Phi- \Phi^\dagger T^a \partial_\mu \Phi ) , $$
$${\mathbb D}_\mu =1\!\! 1 \partial_\mu -ig {\mathbb A}_\mu \equiv 1\!\! 1 \partial_\mu -ig T_aA^a_\mu , \\
{\mathbb D}_\mu \mapsto U {\mathbb D}_\mu U^{-1} , ~~\leadsto \\
{\mathbb A}_\mu \mapsto U {\mathbb A}_\mu U^{-1} -\frac{i}{g} (\partial_\mu U )U^{-1}
$$
the gauge orbit, as amply summarized in texts. The Noether currents will now need to be extended to gauge covariant, instead of the above gauge invariant ones.